Vita
Bruce was born on Sept. 13th, 1950, in
Los Angeles and was educated in Florida and Washington. He received a B.A.
degree at Washington State University in 1976, a M.S. degree from Western
Washington University in 1981 and a Ph.D. from Arizona State University in
1989. Bruce was an enthusiastic member of the American Bryological and
Lichenological Society, The International Association of Lichenology, The
Northwest Lichen Guild, the California Lichen Society and The British
Lichen Society. Through extensive collecting and research, he became one
of the foremost experts in the identification of lichens from the western
U.S. and was a specialist in Lecanora subgenus Placodium. As
the Associate Curator for the Arizona State University Lichen Herbarium,
he was a major contributor to both Vol. I (published 2002) and Vol. II (published
in 2004) of the Lichen Flora of the Greater Sonoran Desert Region.
In the fall of 2002 Bruce was diagnosed with stomach cancer. Although it
his entire stomach was surgically removed the cancer had already spread
and almost a year later cancer was also found in his liver. Intensive
chemotherapy could not stop the cancer and Bruce passed on January 22nd in
Tempe, Arizona.
Afterglow
I'd like the memory of me to be a happy
one,
I'd like to leave an afterglow of smiles when day is gone.
I'd like to leave an echo whispering softly down the ways,
Of happy times, and laughing times, and bright and sunny days.
I'd like the tears of those who grieve, to dry before the sun,
Of happy memories that I leave when life is done
(read at the memorial service for Bruce
Douglas Ryan, January 29, 2004)
Names of "some"
Lichens described by Bruce Ryan
(Note: Some of these names are published in
Volume II of the Sonoran Lichen Flora)
Austropeltis peruviana B.D. Ryan
Cladidium bolanderi (Tuck.) B.D. Ryan
Lecania arizonica B.D. Ryan & Van den Boom
Lecania brattiae B.D. Ryan & T.H. Nash
Lecania caloplacicola B.D. Ryan & Van den Boom
Lecania chalcophila B.D. Ryan & Van den Boom
Lecania fuscelloides B.D. Ryan & Van den Boom
Lecania pacifica Van den Boom & B.D. Ryan
Lecania subdispersa (Nyl. ex B.D. Ryan) B.D. Ryan emend. B.D. Ryan
Lecanora alboloba B.D. Ryan ined.
Lecanora arenisaxicola B.D. Ryan ined.
Lecanora beamanii B.D. Ryan
Lecanora brattiae B.D. Ryan & Van den Boom
Lecanora caesiella B.D. Ryan & T.H. Nash
Lecanora chacoensis B.D. Ryan & T.H. Nash
Lecanora geiserae B.D. Ryan & T.H. Nash
Lecanora heinoi B.D. Ryan
Lecanora kofae B.D. Ryan & T.H. Nash
Lecanora mazatzalensis B.D. Ryan & T.H. Nash
Lecanora muralis var. brunneola (Mereschk.) B.D. Ryan & T.H.
Nash
Lecanora nashii B.D. Ryan
Lecanora pachysoma B.D. Ryan & Poelt
Lecanora phaedrophthalma var. christoi (W.A. Weber) B.D.
Ryan & T.H. Nash
Lecanora pseudomellea B.D. Ryan
Lecanora pueblae B.D. Ryan & T.H. Nash
Lecanora sierrae B.D. Ryan & T.H. Nash
Lecanora sonorae B.D. Ryan & T.H. Nash
Lecanora weberi B.D. Ryan
Lecanora xanthosora B.D. Ryan & Poelt
Lignoscripta atroalba B.D. Ryan
Solenopsora cladonioides B.D. Ryan & Timdal
Xylographa pruinodisca B.D. Ryan & T.H. Nash
Species recently named in honor of Bruce Ryan:
Buellia ryanii Bungartz
Memorial Fund
Bruce's final
lichenological years were devoted to the production of the Greater Sonoran
Desert Lichen Flora and the final volumes will be dedicated to Bruce. As a
way to honor his great contribution to lichenology a memorial fund was
established for the production of color photographs in Vol. III of the
Sonoran Flora. For people with US accounts, checks can be made out to
Arizona State University and sent to Thomas H. Nash III. For those of
you in Europe please do not hesitate to contact
tom.nash@asu.edu.
Publications
This only represents a selection of Bruce's
publications...
Czeczuga, B,
Ryan, BD, Spjut, RW,
Flock, AW, Weber, WA, Beasley, CW, Showman, RE, Worthington, RD, Boucher,
VL 1997: Carotenoids in lichens from the United States of America and
Mexico. - Feddes Repertorium 108(5-6): 401-417.
Goward, T, Breuss, O, Ryan, B, McCune, B,
Sipman, H, Scheidegger, C 1996: Notes on the lichens and allied fungi of
British Columbia. III. - The Bryologist 99(4): 439-449.
Grube, M,
Ryan, BD 2002:
Collemopsidium. - In: Nash, TH, III,Ryan,
BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the Greater
Sonoran Desert Region. I. Lichens Unlimited, Arizona State University,
Tempe, Arizona, pp. 162-164.
Hafellner, J, Triebel, D,
Ryan, BD, Nash, TH, III 2002: On lichenicolous fungi from
North America. II. - Mycotaxon 84: 293-329.
Link, SO,
Ryan, BD, Downs, JL,
Cadwell, LL, Soll, JA, Hawke, MA, Ponzetti, J 2000: Lichens and mosses on
shrub-steppe soils in southeastern Washington. - Northwest Science 74(1):
50-56.
Matthes, U,
Ryan, BD, Larson, DW
2000: Community structure of epilithic lichens on the cliffs of the
Niagara Escarpment, Ontario, Canada. - Plant Ecology 148: 233-244.
Nash, TH, III,
Ryan, BD 2002: Omphalora.
- In: Nash, TH, III,Ryan, BD,Gries,
C,Bungartz, F (eds.): Lichen Flora of the Greater Sonoran Desert Region.
I. Lichens Unlimited, Arizona State University, Tempe, Arizona, pp.
301-302.
Nash, TH, III,
Ryan, BD, Davis, WC,
Breuss, O, Hafellner, J, Lumbsch, HT, Tibell, L, Feuerer, T 1998:
Additions to the lichen flora of Arizona IV. - The Bryologist 101(1):
93-99.
Nash, TH, III,
Ryan, BD, Elix, JA 2002:
Everniastrum. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 191-193.
Nash, TH, III,
Ryan, BD, Gries, C,
Bungartz, F (eds.) 2002: Lichen Flora of the Greater Sonoran Desert
Region. - Lichens Unlimited, Arizona State University, Tempe, Arizona. 532
pp.
Nash, TH, III, Schramm, RT, Herrera, MA,
Ryan, BD 1999: Lichenes
Exsiccati, Distributed by Arizona State University, Fasc. 13 (No.
301-325), Fasc. 14 (No. 326-350). - Tempe, Arizona. 10 pp.
Palmer, FE, Staley, JT, RYAN,
BD 1990: Ecophysiology of microcolonial
fungi and lichens on rocks in northeastern Oregon. - New Phytologist 116:
613-620.
Riefner, RE, Bowler, PA,
Ryan, BD 1995: New and interesting records of lichens from
California. - Bulletin of the California Lichen Society 2(2): 1-11.
RYAN, BD
1989: Lecanora sect. Petrasterion (lichenized Ascomycotina)
in North America: Lecanora weberi Ryan, sp. nov. (subsect.
Pseudocorticatae), from Colorado. - Mycotaxon 36(1): 9-14.
Ryan, BD 1985: Lichens of Chowder Ridge, Mt. Baker, Washington.
- Northwest Science 59: 279-293.
Ryan, BD 1985: New records of lichens from Washington state. -
Evansia 2: 33-48.
Ryan, BD 1988: Marine and maritime lichens on serpentine rock
on Fidalgo Island, Washington. - The Bryologist 91(3): 186-190.
Ryan, BD 1988: Zonation of lichens on a rocky seashore on
Fidalgo Island, Washington. - The Bryologist 91(3): 167-180.
Ryan, BD 1989: A monograph of Lecanora subgen.
Placodium sect. Endochloris (lichenized Ascomycotina). - The
Bryologist 92(4): 513-522.
Ryan, BD 1989: A new species of Lecanora sect.
Dactylon (lichenized Ascomycotina) from Mexico, with notes on other
species of the section in North America. - Cryptogamic Botany 1: 243-248.
Ryan, BD 1990: A study of Lecanora valesiaca (lichenized
Ascomycotina), with notes on L. dispersoareolata, L. albula,
and Squamarina subg. Petroplaca in North America. - Nova
Hedwigia 50(1-2): 81-96.
Ryan, BD 1992: [Review:] P.M. McCarthy. Bibliography of
Australian Lichens: 1807-1991. x + 62 pages. National Herbarium of
Victoria, Melbourne. 1992. - The Bryologist 95(4): 488.
Ryan, BD 1998: A monograph of Lecanora subg.
Placodium sect. Arctoxanthae (lichenized Ascomycotina). - In:
Glenn, MG,Harris, RC,Dirig, R,Cole, MS (eds.): Lichenographia Thomsoniana:
North American Lichenology in Honor of John W. Thomson. Mycotaxon Ltd.,
Ithaca, New York, pp. 105-131.
Ryan, BD 2002: Ahtiana. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 91-92.
Ryan, BD 2002: Alectoria. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 92-95.
Ryan, BD 2002: Baeomyces. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 106-107.
Ryan, BD 2002: Brodoa. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 107-108.
Ryan, BD 2002: Bryoria. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 109-114.
Ryan, BD 2002: Dictyonema. - In: Nash, TH, III,Ryan,
BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the Greater
Sonoran Desert Region. I. Lichens Unlimited, Arizona State University,
Tempe, Arizona, pp. 169-171.
Ryan, BD 2002: Evernia. - In: Nash, TH, III,Ryan,
BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the Greater
Sonoran Desert Region. I. Lichens Unlimited, Arizona State University,
Tempe, Arizona, pp. 188-190.
Ryan, BD 2002: Glypholecia. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 203-204.
Ryan, BD 2002: Imshaugia. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 252-254.
Ryan, BD 2002: Leprocaulon. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 262-264.
Ryan, BD 2002: Letharia. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 267-270.
Ryan, BD 2002: Nodobryoria. - In: Nash, TH, III,Ryan,
BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the Greater
Sonoran Desert Region. I. Lichens Unlimited, Arizona State University,
Tempe, Arizona, pp. 299-301.
Ryan, BD 2002: Parmeliopsis. - In: Nash, TH, III,Ryan,
BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the Greater
Sonoran Desert Region. I. Lichens Unlimited, Arizona State University,
Tempe, Arizona, pp. 315-318.
Ryan, BD 2002: Platismatia. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 400-401.
Ryan, BD 2002: Pseudephebe. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 409-411.
Ryan, BD 2002: Rhizoplaca. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 442-448.
Ryan, BD 2002: Squamarina. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 466-468.
Ryan, BD 2002: Stereocaulon. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 472-474.
Ryan, BD 2002: Vulpicida. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 505-507.
Ryan, BD, Galloway, DJ 2002: Pseudocyphellaria. - In:
Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 413-415.
Ryan, BD, Kalb, K 2002: Dibaeis. - In: Nash, TH, III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of
the Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 168-169.
Ryan, BD, McWhorter, FP 1986: Processing lichen colonies
growing on soil or moss, with glue to facilitate sectioning. - Evansia 3:
14-16.
Ryan, BD, Nash III, TH 1997: Placodioid taxa of Lecanoraceae
sensu Zahlbr. (lichenized Ascomycotina) in North America: taxa excluded
from Lecanora subg. Placodium. - Nova Hedwigia 64(3-4): 393-420.
Ryan, BD, Nash, TH", III 1989: Lecanora sect.
Petrasterion (lichenized Ascomycotina) in North America: Lecanora
sonorae Ryan & Nash, sp. nov. (subsect. Pseudocorticatae), from Mexico.
- Cryptogamic Botany 1: 215-218.
Ryan, BD, Nash, TH, III 1991: Lecanora Sect.
Petrasterion (lichenized Ascomycotina) in North America: Notes on the
L. novomexicana complex (Subsect. Pseudocorticatae). - Mycotaxon
41(1): 57-65.
Ryan, BD, Nash, TH, III 1991: Lichen Flora of the eastern Brook
Lakes Watershed, Sierra Nevada Mountains, California. - The Bryologist
94(2): 181-195.
Ryan, BD, Nash, TH, III 1993: Lecanora sect.
Placodium (Lichenized Ascomycotina) in North America. - Cryptogamic
Botany 3: 264-269.
Ryan, BD, Nash, TH, III 1993: Lecanora section
Placodium (lichenized Ascomycotina) in North America: new taxa in the
L. garovaglii group. - The Bryologist 96(3): 288-298.
Ryan, BD, Nash, TH, III 1995: Lecanora sect.
Petrasterion (lichenized Ascomycotina) in North America: Lecanora
pueblae sp. nov., from Mexico and Lecanora kofae sp. nov., from Arizona. -
Mycotaxon 53: 479-488.
Ryan, BD, Nash, TH, III 1997: Systematics of Lecanora
subgenus Placodium (lichenized Ascomycotina) in North America: an
overview, with keys. - Nova Hedwigia 64(1-2): 111-127.
Ryan, BD, Nash, TH, III 2002: Cetrelia. - In: Nash, TH,
III,
Ryan, BD,Gries, C,Bungartz, F (eds.): Lichen Flora of the
Greater Sonoran Desert Region. I. Lichens Unlimited, Arizona State
University, Tempe, Arizona, pp. 129-130.
Ryan, BD, Nash, TH, III, Herrera-Campos, MA, Hafellner, J,
Lumbsch, HT, Moberg, R, Tibell, L, Ahti, T, Sipman, HJM, Breuss, O 2000:
New records of lichens from M?ico. - Nova Hedwigia 70(1-2): 79-106.
Ryan, BD, Schultz, M 2002: Polychidium. - In: Nash, TH,
III,Ryan, BD,Gries,
C,Bungartz, F (eds.): Lichen Flora of the Greater Sonoran Desert Region.
I. Lichens Unlimited, Arizona State University, Tempe, Arizona, pp.
401-402.
Ryan, BD, Timdal, E 2002: Solenopsora. - In: Nash, TH,
III,Ryan, BD,Gries,
C,Bungartz, F (eds.): Lichen Flora of the Greater Sonoran Desert Region.
I. Lichens Unlimited, Arizona State University, Tempe, Arizona, pp.
462-465.
Smith, C, Geiser, L, Gough, L, McCune, B,
RYAN, BD, Showman,
R 1993: Species and communities. - In: Stolte, K,Mangis, D,Doty,
R,Tonnessen, K,Huckaby, LS (eds.): Lichens as Bioindicators of Air Quality.
General Technical Report, U.S. Department of Agriculture, Forest Service,
Rocky Mountain Forest and Range Experiment Station, Fort Collins,
Colorado, pp. 41-66. |
