vascular plant collections from southwestern United States and northern
- Especially important collections
voucher specimens of vascular plants
- Extensive collections of certain genera of Asteraceae
- A large collection of American Myrtaceae
collections from temperate South America, especially Chile
Most of the growth of the ASU Vascular Plant Herbarium has occurred
in the last 25 years. Specimens have accumulated through the activities
of Arizona State University staff, graduate students, and others associated
with the herbarium. Our areas of most intense study, the Southwest and
northern Mexico, are areas in which much botanical exploration is underway
and still needed. This need is evidenced by the number of new state records
and newly described taxa that have been based on our collections. With
funds provided by a National Science Foundation grant to Noel Holmgren
at the New York Botanical Garden, extensive collections (ca. 14,000) were
made throughout the southwestern United States during the years 1973 to
1975 by D. J. Pinkava, E. Lehto, and D. J. Keil. Aided by another National
Science Foundation grant to W. L. Minckley, D. J. Pinkava also conducted
a floristic study of the Cuatro Cienegas region of Coahuila, Mexico (Pinkava
The following graduate students of D. J. Pinkava and Les Landrum (as committee chair,
co-chair, or member) have done floristic or related studies for master's
degrees in various parts of Arizona and have added their collections to
the herbarium (chronologically): E. Lehto (Lake Pleasant Regional Park,
Maricopa Co.); D. J. Keil (White Tank Mountains Regional Park, Maricopa
Co.); M. A. Lane (McDowell Mountain Regional Park, Maricopa Co.); E. G.
Sundell (Sierra Estrella Regional Park, Maricopa Co.); S. Forbes, (Pinal
Mountains, Gila Co.); M. Russo (Castle Dome Mountains, Yuma Co.); T. Reeves
(Chiricahua National Monument, Cochise Co.); J. Leithliter (Chiricahua
National Wilderness Area, Cochise Co.); G. Marrs-Smith (San Bernadino Ranch,
Cochise Co.); W. Johnson (Pinaleno Mountains, Graham Co.); A. Pierce (Buckeye
Hills Semi-Regional Park, Maricopa Co.); K. Rice (Superstition Wilderness
Area); L. Wolden (Hassayampa River Reserve); G. Imdorf (Sierra Ancha Wilderness
Area); J. Buegge (Santa Theresa Mountains, Graham Co.); D. Damrel (ASU Arboretum); S. Doan (Seven Springs Recreation Area, Maricopa Co.); E. Gilbert,
(West Fork Oak Creek Canyon, Coconino Co.); E. Makings (San Pedro Riparian National Conservation Area, Cochise Co.).,
B. Lester (Hummingbird Springs Willderness Area, Maricopa Co.).
Other important collections made by persons who deposited their specimens
at the Arizona State University Herbarium but for non-degree studies are:
B. Walden (Usery Mountain Park, Maricopa Co.); E. Lehto (Cabeza Prieta
Game Range, Yuma, Co.); W. L. Minckley and L. A. McGill (upper reaches
of Gila River); R. K. Gierisch, G. K. Brown and B. D. Parfitt (northwestern
Arizona north of the Colorado, River, i.e., the Arizona Strip); M. Butterwick
and T. Daniel (South Mountain Park, Maricopa Co.); M. Butterwick and B.
D. Parfitt (Hualapai Mountains, Mohave Co.); H. Tate (Peña Blanca
Lake area, Santa Cruz Co.); M. Butterwick et al. (Hualapai-Aquarius, Harcuvar,
Vulture, and Skull Valley Planning Units); R. L. Burgess (Tonto National
Monument, Gila Co.); M. Baker and T. Wright (Camp Wood, Williamson Valley,
Yolo North, and Yolo South grazing allotments on the Chino Valley Ranger
District, USFS); M. Baker and T. Wright (Apache Creek, Juniper Mesa, Sycamore
Canyon, and Woodchute Wilderness areas of Prescott National Forest); J.
R. Sutherland (Show Low Lake area); B. D. Parfitt and C. Christy (Coronado
National Monument). Most, if not all, of the collections listed above are
the most complete in existence for these particular areas. Through an active
exchange program with other institutions in the Southwest and Mexico we
have enriched our regional collection significantly.
2) Collections of Cactaceae
One of the centers of diversity of the Cactaceae is the southwestern
United States and northern Mexico. It is logical then that our collection
of ca. 5,000 specimens of this family be outstanding. Because cacti are
not easily collected, there are few institutions that have good fertile
collections. The ASU Herbarium Cactaceae collection is perhaps among the top five in the
United States surpassed only by MEXU (Mexico City) outside the United
States in the Western Hemisphere. In addition, ASU has the best collection
of cytological voucher specimens of Cactaceae in the world. Many of the
specimens are accompanied by photographs of the plants in their natural
habitat, and close-ups of important structures. ASU's holdings of Cactaceae
are enhanced by degree studies: on hybridization done on the following
taxa: x Myrtgerocactus (L. A. McGill); Opuntia x kelvinensis
(M. A. Baker); O. curvospina (B. D. Parfitt); and the O.whipplei
complex (N. Trushell); fleshy-fruited prickly-pears Opuntia subgen.
Opuntia in Arizona (M. G. McLeod); the O. polyacantha complex
in western North America (B. Parfitt); the Echinocactus polycephalus
complex (M. Chamberland); and Opuntia subgen. Cylindropuntia
in Baja California (J. Rebman).
voucher specimens of vascular plants
Since D. J. Pinkava came to Arizona State University the ASU Herbarium
has been a center for cytological and hybridization studies of Southwestern
plants. In addition to Pinkava, several of his students have contributed
to cytological studies at ASU, most notably R. C. Brown, D. J. Keil, M.
A. Baker, B. D. Parfitt, T. Reeves, G. K. Brown, J. P. Rebman, and C. Christy.
Over the years 35 publications on chromosome counts listing more than 1700
voucher specimens at ASU have been produced. In many cases we have been
able to exchange with other institutions interested in cytological studies
(e.g., NMS, OS, UC, TEX) and thus have many additional cytological vouchers
in our collection.
The Asteraceae at ASU are important because of revisionary or monographic
studies on following genera conducted by staff and graduate students: Psilostrophe
(R. C. Brown, 1977, 1978) and Baileya (R. C. Brown and Pinkava,
1974), Platyschkuhria (G. K. Brown, 1983) and Haplopappus
of South America (G. K. Brown and Clark, 1982), Tragopogon of Arizona
(R. K. Brown, 1974 unpublished thesis), Berlandiera (Pinkava,1967),
Hazardia (Clark1979), and Hymenothrix (Keil and Pinkava,
L. R. Landrum is a specialist in American Myrtaceae and receives numerous
gifts for determination and collects Myrtaceae for his own studies [Myrtaceae
of Chile (1988b), Myrteola (1988a), Chamguava (1991), Mosiera
(1992), Myrtinae of Mesoamerica, Psidium and
related genera (2003, 2005), such that ASU now has a collection of ca. 11,000
American Myrtaceae. It is probably one of the larger university collections
of Myrtaceae in the United States.
L. R. Landrum is participating in the Flora of Chile project, providing
treatments of the Berberidaceae and Myrtaceae, and has made numerous trips
to Brazil, Chile and Argentina. Thousands of collections
have been made and exchanges have been established with MBM, CTES, HUEFS, MO,and
other herbaria in order to increase ASU's collection of temperate South American collections.
Few herbaria outside of Chile have significant Chilean collections. The ASU collection of Chilean
Berberis is probably the best outside of Chile.
45% - ARIZONA
25% - REST OF UNITED STATES AND CANADA
25% - AMERICAS SOUTH OF THE UNITED STATES
5% - OLD WORLD AND CULTIVATED
Brown, G.K. 1983. Chromosome numbers in Platyschkuhria Rydberg
(Compositae) and their systematic significance. American Journal of Botany
Brown, G.K. and W.D. Clark. 1982. Taxonomy of Haplopappus section
Gymnocoma (Compositae). Systematic Botany 7: 199-213.
Brown, R.C. 1978. Biosystematics of Psilostrophe DC. (Compositae).
I. Chromosome variability. Rhodora 79: 169-189.
Brown, R.C. 1978. Biosystematics of Psilostrophe (Compositae:
Helenieae). II. Artificial hybridization and systematic treatment. Madroño
Brown, R.C. and D.J. Pinkava. 1974. Distribution and meiotic behavior
of supernumerary chromosomes in Baileya multiradiata (Compositae).
Bulletin of the Torrey Botanical Club 101: 177-181.
Clark, W.D. 1979. The taxonomy of Hazardia (Compositae: Asteraceae).
Madroño 26: 105-127.
Kearney, T.H., R.H. Peebles and collaborators. 1942. Flowering Plants
and Ferns of Arizona. U. S. Department of Agriculture Miscellaneous Publication
No. 428, Washington, D. C.
Kearney, T.H., R.H. Peebles and collaborators. 1951. Arizona Flora.
University of California Press: Berkeley.
Kearney, T.H., R.H. Peebles and collaborators. 1960. Arizona Flora.
University of California Press: Berkeley. Ed. 2 with Supplement by J. T.
Howell, E. McClintock and collaborators.
Landrum, L. R. 1988a. Systematics of Myrteola (Myrtaceae). Systematic
Botany 13(1): 120-132.
Landrum, L. R. 1988b. The Myrtle Family (Myrtaceae) in Chile. Proc.
Calif. Acad. Sciences 45(12): 277-317.
Landrum, L. R. 1991. Chamguava: a new genus of Myrtaceae (Myrtinae)
from Mesoamerica. Systematic Botany 16(1): 21-29.
Landrum, L. R. 1992. Mosiera (Myrtaceae) in Mexico and Mesoamerica.
Novon 2: 26-29.
Landrum, L. R. Myrtinae (Myrtaceae) of Mesoamerica.
35 manuscript pages. Submitted to the Missouri Botanical Garden, Flora
of Mesoamerica project.
Landrum, L. R. 2005. A revision of the Psidium grandifolium complex (Myrtaceae). Sida 21(3): 1335-1354. 2005.
Ladnrum, L. R. 2003. A revision of the Psidium salutare complex (Myrtaceae). Sida 20(4): 1449-1469. 2003.
Lehr, J.H. 1978. A catalogue of the flora of Arizona. Desert Botanical
Lehr, J. H. and D. J. Pinkava. 1980. A catalogue of the Flora of Arizona.
Supplement I. Journal of the Arizona-Nevada Academy of Science 15: 17-32.
Lehr, J. H. and D. J. Pinkava. 1982. A catalogue of the Flora of Arizona.
Supplement II. Journal of the Arizona-Nevada Academy of Science 17: 19-26.
Pinkava, D. J. 1967. Biosystematic study of Berlandiera (Compositae).
Brittonia 19: 285-298.
Tidestrom, I. and T. Kittle. 1941. A Flora of Arizona and New Mexico.
The Catholic University Press, Washington, D. C.